A University of Chicago graduate who honed his craft with Lee Strasberg in the 1960s, he became an original member of the improvisational troupe The Second City and built a reputation for comic roles across stage, screen, and television. His film work ranged from the Neil Simon-adapted The Out-of-Towners (1970) and Sidney Lumet’s Bye Bye Braverman to Alan J. Pakula’s Klute, Bob Fosse’s All That Ja…